The true finches
are passerine birds in the family Fringillidae
. They are predominantly seed-eating songbirds. Most are native to Southern Hemisphere, but one subfamily is endemic to the Neotropics, one to the Hawaiian Islands, and one subfamily – monotypic at genus level – is found only in the Palaearctic. The scientific name Fringillidae comes from the Latin word fringilla
for the Chaffinch (Fringilla coelebs
) – a member of that last subfamily – which is common in Europe.
Many birds in other families are also commonly called "finches", including some species in the very similar-looking waxbills (family Estrildidae) of the Old World tropics and Australia; several groups of the bunting and American sparrow family (Emberizidae); and Darwin's finches of the Galapagos islands, which provided evidence of natural selection and are now recognized to be peculiar tanagers (Thraupidae). [1]
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FINCH TICKETS
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Description
thumb
The "classical" true finches range in size from the
Andean Siskin (
Carduelis spinescens
) at 9.5 cm (3.8 in) and 8.4 g (0.3 oz) to the
Collared Grosbeak (
Mycerobas affinis
) with its nearly 23 cm (9 in) and 80 g (3 oz). They typically have strong, stubby
beaks, which in some species can be quite large; however,
Hawaiian honeycreepers are famous for the wide range of bill shapes and sizes brought about by
adaptive radiation. All true finches have 12
remiges and 9 primary
rectrices. The basic
plumage colour is brownish, sometimes greenish; many have considerable amounts of black, while white plumage is generally absent except as wing-bars or other signalling marks. Bright yellow and red
carotenoid pigments are commonplace in this family, and thus blue
structural colours are rather rare, as the yellow pigments turn the blue color into green. Many, but by no means all true finches have strong
sexual dichromatism, the females typically lacking the bright carotenoid markings of males.
Finches are typically inhabitants of well-wooded areas, but some can be found on mountains or even in
deserts. They are primarily
granivorous, but
euphoniines include considerable amounts of
arthropods and
berries in their diet, and Hawaiian honeycreepers
evolved to utilize a wide range of food sources, including
nectar. The diet of Fringillidae
nestlings includes a varying amount of small arthropods. True finches have a bouncing flight like most small
passerines, alternating bouts of flapping with gliding on closed wings. Most sing well and several are commonly-seen
cagebirds; foremost among these is the
domesticated Canary (
Serinus canaria domestica
). The nests are basket-shaped and usually built in trees, more rarely in bushes, between rocks or on similar substrate.
Systematics and taxonomy
The taxonomic structure of the true finch family, Fringillidae, has been fairly disputed in the past, with some upranking the
Hawaiian honeycreepers (Drepanidinae) as family Drepanididae and/or uniting the cardueline and fringilline finches as
tribes (Carduelini and Fringillini) in one subfamily; the euphonious finches (
Euphoniinae) were thought to be
tanagers due to general similarity in appearance and mode of life until their real affinities were realized. In particular
North American authors have often have merged the
buntings and
American sparrow family (
Emberizidae) – and sometimes the bulk of the
nine-primaried oscines – with the split-up Fringillidae as subfamilies of a single massive family. But the current understanding of
Passeroidea phylogeny is better reflected in keeping the fundamental nine-primaried oscine
clades as distinct families. However,
Przewalski's "Rosefinch" (
Urocynchramus pylzowi
) is now classified as a distinct family, monotypic as to genus and species, and with no particularly close relatives among the Passeroidea.
[2]
Fossil remains of true finches are rare, and those that are known can mostly be assigned to extant
genera at least. Like the other Passeroidea families, the true finches seem to be of roughly
Middle Miocene origin, around 20-10
million years ago (Ma). An unidentifable finch fossil from the
Messinian age, around 12 to 7.3
million years ago (Ma) during the
Late Miocene subepoch, has been found at
Polgárdi in
Hungary.
[3]
Subfamilies and genera
thumb (
Coccothraustes coccothraustes
), one of the
Holarctic grosbeaks
thumb (
"Carpodacus" cassinii
) is not part of the genus
Carpodacus
proper
thumb (
Carpodacus roseus
), a true rosefinch
thumb (
Carduelis (Spinus) magellanica
)
thumb (
Vestiaria coccinea
), a
Hawaiian honeycreeper
thumb (
Euphonia violacea
)
The systematics of the cardueline finches are contentious. The layout presented here follows the recent decades'
molecular phylogenetic studies, and takes into account the traditional splitting of the genus
Carduelis
. The exact position of several genera in the cardueline sequence is tentative.
[4]
- Subfamily Fringillinae
– fringilline finches
. Three species, which feed their young on insects and few if any seed
- * Genus Fringilla
- Subfamily Carduelinae
– cardueline finches
. A much larger group that contains several genera which feed their young mainly on seeds. This subfamily consists of several well-defined clades.
- * Typical grosbeaks
- ** Genus Eophona
– Oriental grosbeaks
- ** Genus Coccothraustes
– Holarctic grosbeaks (polyphyletic?)
- ** Genus Mycerobas
– Mycerobas grosbeaks
- * American rosefinches
- ** Genus "Carpodacus"
– House Finch, Cassin's Finch, and Purple Finch
- * Bullfinches
- ** Genus Pinicola
– Pine Grosbeak
- ** Genus Pyrrhula
– bullfinches
- * Arid-zone clade
- ** Genus Leucosticte
– mountain finches
- ** Genus N.N. – Dark-breasted Rosefinch, "Carpodacus" nipalensis
- ** Genus Rhodopechys
– Trumpeter Finch and relatives
- * Asian rosefinches
- ** Genus Carpodacus
– rosefinches (Genus in need of substantial revision. May consist of two or more genera; alternatively may include Haematospiza
and Uragus
)
- ** Genus Haematospiza
– Scarlet Finch
- ** Genus Uragus
– streaked rosefinches
- *Goldfinch-canary-crossbill clade
(Carduelis
and Serinus
are polyphyletic and are probably best regarded as a collection of independent sub-groups or genera.)
- ** Genus Serinus sensu lato
– canaries, African seedeaters, serins and African siskins
- *** (Sub)Genus Serinus sensu stricto
– European Serin and relatives (possibly related to Spinus
)
- *** (Sub)Genus Crithagra
– Afrotropical canaries, African seedeaters, citrils (probably only distantly related to Serinus
)
- ** Genus Linurgus
– Oriole Finch (possibly basal to Crithagra
)
- ** Genus Rhynchostruthus
– golden-winged grosbeaks (tentatively placed here)
- ** Genus "Serinus" thibetanus
– Tibetan Serin (probably related to Spinus
and Serinus sensu stricto
)
- ** Genus Carduelis sensu lato
- *** (Sub)Genus Carduelis sensu stricto
– European Goldfinch, Citril Finch and Corsican Finch (probably only distantly related to Spinus
)
- *** (Sub)Genus Spinus
– siskins and American goldfinches
- *** (Sub)Genus Linaria
– linnets and twite (related to Spinus
)
- *** (Sub)Genus Chloris
– greenfinches and Desert Finch
- *** (Sub)Genus Acanthis
– redpolls (related to Loxia
)
- *** (Sub)Genus Loxia
– crossbills
- * Carduelinae
incertae sedis
- ** Genus Pyrrhoplectes
– Gold-naped Finch
- ** Genus Chaunoproctus
– Bonin Grosbeak (extinct: 1830s)
- ** Genus Callacanthis
– Spectacled Finch
- ** Genus Neospiza
– Sao Tomé Grosbeak
- Subfamily Drepanidinae
– Hawaiian honeycreepers
. Endemic to the Hawaiian Islands; formerly often treated as a separate family.
- * Some 10-12 living genera, about 7 recently extinct
- Subfamily Euphoniinae
– Euphonious finches
. Endemic to the Neotropics; formerly placed in Thraupidae.
- * Genus Euphonia
– euphonias
- * Genus Chlorophonia
– chlorophonias
Footnotes
- Newton (1973), Clement ''et al.'' (1993)
- Clement ''et al.'' (1993), Groth (2000), Jønsson & Fjeldså (2006), Arnaiz-Villena ''et al.'' (2007)
- Hír ''et al.'' (2001), Mlíkovský (2002)
- Marten & Johnson (1986), Arnaiz-Villena ''et al.'' (1998, 2001, 2007, 2008)
References
- Newton (1973), Clement ''et al.'' (1993)
- Clement ''et al.'' (1993), Groth (2000), Jønsson & Fjeldså (2006), Arnaiz-Villena ''et al.'' (2007)
- Hír ''et al.'' (2001), Mlíkovský (2002)
- Marten & Johnson (1986), Arnaiz-Villena ''et al.'' (1998, 2001, 2007, 2008)